Lust is a Homunculus and one of the first antagonists in the Fullmetal Alchemist series. Lust is the first Homunculus to appear in the series and plays a significant part in the plots of both the manga and 2003 anime series, usually partnered with Gluttony to whom she appears to have a particular, almost motherly closeness. Her Ouroboros mark is located on her collarbone, just above her breasts.